Students entering through any of the eight themes in the Graduate Biomedical Sciences will have several common courses throughout their first year, including basic biosciences, biostatistics, and ethics. Intermingled with these common courses will be theme-specific courses. Lab rotations (3) are unique to the individual, although everyone's rotations begin and end at the same time. Click here for an overview.


For the students entering in Fall 2014, the tentative lab rotation schedule is as follows:

  • 1st Lab Rotation - 25 August through 19 November 2014
         Poster Session - 19 November 2014
  • 2nd Lab Rotation - 20 November 2014 through 27 February 2015
         Poster Session - 26 February 2015
  • 3rd Lab Rotation - 2 March 2015 through 27 May 2015
         Poster Session - 27 May 2015

NOTES: Students making a C or lower (considered a failure) in any course must retake and pass the course the following year. Any student receiving two C's (or more) are subject to being dismissed by the program.


All students in the GBS themes take three basic core courses, which meet M-F, 8 a.m.-10 a.m., in BBRB 170:

GBS 707 - Basic Biochemistry and Metabolism
12 August 2014 - 15 September 2014   

  • Amino acids and primary protein structure
  • Protein secondary and tertiary structure
  • Postranslational modifications  and allosteric changes
  • Glycobiology
  • Lipid structure and metabolism
  • Thermodynamic principles of biochemical reactions
  • Enzyme kinetics
  • Enzyme mechanisms and regulation
  • Amino acid/nitrogen metabolism
  • Glycolysis, gluconeogenesis, and citric acid cycle
  • Glycogenesis, glycogenolysis, and pentose phosphate pathway
  • Electron transport/oxidative phosphorylation  

GBS 708 - Basic Genetics and Molecular Biology
16 September 2014 - 20 October 2014

  • DNA recombination
  • Nucleic acids & DNA replication
  • Prokaryotic transcription, translation, and gene regulation 
  • Eukaryotic genome organization
  • Eukaryotic DNA replication
  • Eukaryotic transcription, translation, and gene regulation
  • Mendelian inheritance
  • Genetic variation and polymorphisms

After these courses, students break into their theme-specific coursework for the remainder of the Fall semester. Neuroscience students take:

  • GBS 731 - Principles of Cellular Neuroscience (M-F, 8 a.m.-10 a.m., 21 October - 22 December 2014)
  • GBS 734 - Neuroscience Historical Literature Discussion Group (September-December, 2014)


SPRING SEMESTER 2015 (Dates below for 2015 are tentative)

5 January - 27 February
     GBS 732 - Grad Neuroscience
          This course is taken with dental and optometry students and includes a discussion group component. 
          Course meets Mondays & Wednesdays, 1 p.m.-2:50 p.m., and Fridays, 8 a.m.-11:50 a.m.;
           the discussion group meets Mondays & Wednesdays, 3:30pm-5:30pm.

3 March - 30 March
     GBS 714 - Developmental Neuroscience
          Meets Monday-Friday, 8 a.m.-10 a.m.

31 March - 27 April
     GBS 733 - Diseases of the Nervous System
          Meets Monday-Friday, 8 a.m.-10 a.m.



Neuroscience students will also attend the Student Summer Seminar Series (GBS 737), each Thursday at 1:30 p.m., May-September. In this seminar series, (for 2014 the dates were May 8-July 24) advanced neuroscience students from across campus present their research work in a professional seinar setting. Each week, two students present to faculty and fellow students to gain valuable experience in this important component of scientific work.


GRD 717 - Bioethics (required for all GBS students beginning their 2nd year; meets Friday afternoons)

BY 755 - Biometry (stats course that meets Tuesday and Thursday afternoons, 2-3pm)

Other advanced program academic requirements:

    • Seminar every semester (GBS 737 every summer semester)
    • Journal club every semester
    • 3 advanced courses chosen by the student and permanent mentor
    • Year-End Assessment (written and oral) in the summer of first year
    • Thesis committee formed early during second year
    • Defense of thesis proposal between October and March of student's third year (official Qualifying Exam)